What does Albert Einstein mean by the word “mollusc” and how does it fit in his theory of Relativity? The word can be found in chapter 29 of Relativity: The Special and General Theory.
In gravitational fields there are no such things as rigid bodies with Euclidean properties; thus the fictitious rigid body of reference is of no avail in the general theory of relativity. The motion of clocks is also influenced by gravitational fields, and in such a way that a physical definition of time which is made directly with the aid of clocks has by no means the same degree of plausibility as in the special theory of relativity.
For this reason non-rigid reference-bodies are used, which are as a whole not only moving in any way whatsoever, but which also suffer alterations in form ad lib. during their motion. Clocks, for which the law of motion is of any kind, however irregular, serve for the definition of time. We have to imagine each of these clocks fixed at a point on the non-rigid reference-body. These clocks satisfy only the one condition, that the “readings” which are observed simultaneously on adjacent clocks (in space) differ from each other by an indefinitely small amount. This non-rigid reference-body, which might appropriately be termed a “reference-mollusc”, is in the main equivalent to a Gaussian four-dimensional co-ordinate system chosen arbitrarily. That which gives the “mollusc” a certain comprehensibility as compared with the Gauss co-ordinate system is the (really unjustified) formal retention of the separate existence of the space co-ordinates as opposed to the time co-ordinate. Every point on the mollusc is treated as a space-point, and every material point which is at rest relatively to it as at rest, so long as the mollusc is considered as reference-body. The general principle of relativity requires that all these molluscs can be used as reference-bodies with equal right and equal success in the formulation of the general laws of nature; the laws themselves must be quite independent of the choice of mollusc.